has been still less in some other classes, as in that
of the Crustacea, for here the most wonderfully diverse
forms are still tied together by a long, but broken,
chain of affinities. Extinction has only separated
groups: it has by no means made them; for if
every form which has ever lived on this earth were
suddenly to reappear, though it would be quite impossible
to give definitions by which each group could be distinguished
from other groups, as all would blend together by
steps as fine as those between the finest existing
varieties, nevertheless a natural classification, or
at least a natural arrangement, would be possible.
We shall see this by turning to the diagram:
the letters, A to L, may represent eleven Silurian
genera, some of which have produced large groups of
modified descendants. Every intermediate link
between these eleven genera and their primordial parent,
and every intermediate link in each branch and sub-branch
of their descendants, may be supposed to be still
alive; and the links to be as fine as those between
the finest varieties. In this case it would be
quite impossible to give any definition by which the
several members of the several groups could be distinguished
from their more immediate parents; or these parents
from their ancient and unknown progenitor. Yet
the natural arrangement in the diagram would still
hold good; and, on the principle of inheritance, all
the forms descended from A, or from I, would have
something in common. In a tree we can specify
this or that branch, though at the actual fork the
two unite and blend together. We could not, as
I have said, define the several groups; but we could
pick out types, or forms, representing most of the
characters of each group, whether large or small,
and thus give a general idea of the value of the differences
between them. This is what we should be driven
to, if we were ever to succeed in collecting all the
forms in any class which have lived throughout all
time and space. We shall certainly never succeed
in making so perfect a collection: nevertheless,
in certain classes, we are tending in this direction;
and Milne Edwards has lately insisted, in an able
paper, on the high importance of looking to types,
whether or not we can separate and define the groups
to which such types belong.
Finally, we have seen that natural selection, which results from the struggle for existence, and which almost inevitably induces extinction and divergence of character in the many descendants from one dominant parent-species, explains that great and universal feature in the affinities of all organic beings, namely, their subordination in group under group. We use the element of descent in classing the individuals of both sexes and of all ages, although having few characters in common, under one species; we use descent in classing acknowledged varieties, however different they may be from their parent; and I believe this element of descent is the hidden bond of connexion which naturalists


