The reindeer-caribou type, of the genus Rangifer, agrees with American deer in having the vertical plate of the vomer complete, and in having the lower ends of the lateral metacarpals remaining, but, like Cervus, it has a brow-tine to the antlers.  Of its early history we know nothing, for the only related forms which have yet come to light are of no great antiquity, being confined to the Pleistocene of Europe as far south as France, and are not distinguishable from existing species.  Until recently it has been supposed that one species was found in northern Europe and Asia, and two others, a northern and a southern, in North America, but lately the last two have been subdivided, and the present practice is to regard the Scandinavian reindeer (Rangifer tarandus) as the type, with eight or nine other species or sub-species, consisting of the two longest known American forms, the northern, or barren-ground caribou (R. arcticus); the southern, or woodland (R. caribou); the three inhabiting respectively Spitzbergen, Greenland and Newfoundland, and still more lately four more from British Columbia and Alaska.  The differences between these are not very profound, but they seem on the whole to represent two types:  the barren-ground, small of size, with long, slender antlers but little palmated; and the woodland, larger, with shorter and more massive antlers, usually with broad palms.  There is some reason to believe that both these types lived in Europe during the interglacial period, the first-named being probably the earlier and confined to western Europe, while the other extended into Asia.  The present reindeer of Greenland and Spitzbergen seem to agree most closely with the barren-ground, while the southern forms are nearest to the woodland, and these are said to also resemble the reindeer of Siberia.  It is, therefore, not an improbable conjecture that there were two migrations into America, one of the barren-ground type from western Europe, by way of the Spitzbergen land connection, and the other of the woodland, from Siberia, by way of Alaska.

Little more can be said, perhaps even less, of the other circumpolar genus, Alces, known in America as “moose,” and across the Atlantic as “elk.”  It also is of mixed character in relation to the two great divisions we have had in mind, but in a different way from reindeer.

Like American deer it has the lower ends of the lateral metacarpals remaining, and the antlers are without a brow-tine, but like Cervus it has an incomplete vomer, and unlike deer in general, the antlers are set laterally on the frontal bone, instead of more or less vertically, and the nasal bones are excessively short.  The animal of northern Europe and Asia is usually considered to be distinct from the American, and lately the Alaskan moose has been christened Alces gigas, marked by greater size, relatively more massive skull, and huge antlers.  Of the antecedents