Water for Elephants

What is the theme in Water for Elephants by Sara Gruen?

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The theme of love is hard to miss, despite all the circus cruelties. Young Jacob falls instantly in love with Marlena, the trick horse rider. Later Marlena works with Rosie, the elephant, with whom both Jacob and Marlena fall in love. It is their love of animals that pulls the twenty-year-olds together.