(Footnote. Mem. de l’Acad. des Scien. de Paris 1775 page 518.)
The second view suggested, in which the anthera in Cycadeae is considered as producing on its surface an indefinite number of pollen masses, each enclosed in its proper membrane, would derive its only support from a few remote analogies: as from those antherae, whose loculi are sub-divided into a definite, or more rarely an indefinite, number of cells, and especially from the structure of the stamina of Viscum album.
I may remark, that the opinion of M. Richard,* who considers these grains, or masses, as unilocular antherae, each of which constitutes a male flower, seems to be attended with nearly equal difficulties.
(Footnote. Dict. Class. d’Hist. Nat. tome 5 page 216.)
The analogy between the male and female organs in Coniferae, the existence of an open ovarium being assumed, is at first sight more apparent than in Cycadeae. In Coniferae, however, the pollen is certainly not naked, but is enclosed in a membrane similar to the lobe of an ordinary anthera. And in those genera in which each squama of the amentum produces two marginal lobes only, as Pinus, Podocarpus, Dacrydium, Salisburia, and Phyllocladus, it nearly resembles the more general form of the antherae in other Phaenogamous plants. But the difficulty occurs in those genera which have an increased number of lobes on each squama, as Agathis and Araucaria, where their number is considerable and apparently indefinite, and more particularly still in Cunninghamia, or Belis,* in which the lobes, though only three in number, agree in this respect, as well