Warburg studied chemistry at the University of Freiburg beginning in 1901. After two years, he left for the University of Berlin to study under Emil Fischer, and in 1906 received a doctorate in chemistry. His interest turned to medicine, particularly to cancer, so he continued his studies at the University of Heidelberg where he earned an M.D. degree in 1911. He remained at Heidelberg, conducting research for several more years and also making several research trips to the Naples Zoological Station.
Warburg's career goal was to make great scientific discoveries, particularly in the field of cancer research, according to the biography written by Hans Adolf Krebs , one of Warburg's students and winner of the 1953 Nobel Prize in medicine and physiology. Although he did not take up problems specifically related to cancer until the 1920s, his early projects provided a foundation for future cancer studies. For example, his first major research project, published in 1908, examined oxygen consumption during growth. In a study using sea urchin eggs, Warburg showed that after fertilization, oxygen consumption in the specimens increased 600 percent.
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